Asahigaoka might look like typical, boring countryside to most; however, no day in this village can ever be considered colorless thanks to five students of varying ages occupying the only class in the only school in town. The youngest student is first grader Renge Miyauchi, who brings an unadulterated wit, curiosity, and her characteristic catchphrase, "Nyanpasu!" Then there are the Koshigaya siblings consisting of the quiet ninth grader and elder brother Suguru, diminutive eighth grader Komari, and the mischievous seventh grader Natsumi. The recent arrival of Tokyo-raised fifth grader Hotaru Ichijou, who appears overdeveloped for her age and thus naturally holds an air of maturity, rounds out this lively and vibrant group of five classmates.

The carefree journeys of the four girls continue! As their fleeting yet eventful summer becomes a distant memory, the arrival of fall signals a new school year with limitless opportunities. Even as the four girls return to their peaceful routine, the appearance of new faces enlivens the anything but exciting rural life of Asahigaoka.